Pilegaard Lab

Peer Reviewed publications

Articles

1. Juel, C, A Honig and H Pilegaard. 1991. Muscle lactate transport studied in sarcolemmal giant vesicles: dependence on fiber type and age. Acta Physiol. Scand. 143: 361-365.

2. Pilegaard, H, C Juel, and F Wibrand. 1993. Lactate transport studied in sarcolemmal giant vesicles from rats: effect of training. Am. J. Physiol. 264: E156-E160.

3. Juel, C, S Kristiansen, H Pilegaard, J Wojtaszewski, and EA Richter. 1994. Kinetics of lactate transport in sarcolemmal giant vesicles obtained from human skeletal muscle. J. Appl. Physiol. 76(3): 1031-1036.

4. Pilegaard, H, J Bangsbo, EA Richter and C Juel. 1994. Lactate transport studied in sarcolemmal giant vesicles from human muscle biopsies: relation to training status. J. Appl. Physiol. 77(4): 1858-1862.

5. Pilegaard, H and C Juel. 1995. Lactate transport studied in sarcolemmal giant vesicles from rat skeletal muscles: effect of denervation. Am. J. Physiol. 269: E679-E682.

6. Pilegaard, H, J Bangsbo, P Henningsen, C Juel, EA Richter. 1995. The effect of blood flow on muscle lactate release studied in the perfused rat hindlimb. Am. J. Physiol. 269: E1044-E1051.

7. Pilegaard, H, T Mohr, M Kjær, C Juel. 1998. Laktat/H+ transport in skeletal muscle rom spinal-cord-injured patients. Scand. J. Med. Science Sports. 8: 98-101.

8. Pilegaard, H and S Asp. 1998. Effect of prior eccentric contractions on lactate/H+ transport in rat skeletal muscle. Am. J. Physiol. 274: E554-E559.

9. Rådegran, G, H Pilegaard, JJ Nielsen, J Bangsbo. 1998. Microdialysis ethanol removal reflects probe recovery rather than local blood flow in skeletal muscle. J. Appl. Physiol. 85(2): 751-757.

10. Wilson, MC, VN Jackson, C Heddle, NT Price, H Pilegaard, C Juel, A Bonen, I Montgomery, O. Hutter, and A.P. Halestrap. 1998. Lactic acid efflux from white skeletal muscle is catalyzed by the monocarboxylate transporter isoform MCT3. J. Biol. Chem. 273(26): 15920-15926.

11. Juel, C and H Pilegaard. 1998. Lactate/H+ transport kinetics in rat skeletal muscle related to fibre type and changes in transport capacity. Pflügers Arch. - Eur. J. Physiol. 436: 560-564.

12. Pilegaard, H, K Domino, T Noland, C Juel, Y Hellsten, B Saltin, J Bangsbo. 1999. Effect of high intensity training on lactate/H+ transport in human skeletal muscle. Am. J. Physiol. 276: E255-E261.

13. Pilegaard, H G Terzis, A Halestrap, and C Juel. 1999. Distribution of the lactate/H+ transporter isoforms MCT1 and MCT4 in human skeletal muscle. Am. J. Physiol. 276: E843-E848.

14. Juel, C H Pilegaard, JJ Nielsen, and J Bangsbo. 1999. Interstitial K+ in human skeletal muscle during and after dynamic graded exercise determined by microdialysis. Am. J. Physiol. 278: R400-406.

15. Pilegaard, H, GA Ordway, B Saltin, PD Neufer. 2000. Transcriptional regulation of gene expression in human skeletal muscle during recovery from exercise. Am. J. Physiol. 279:E806-E814.

16. Keller, C, A Steensberg, H Pilegaard, T Osada, B Saltin, B Klarlund Petersen, and PD Neufer. 2001. Transcriptional Activation of the IL-6 Gene in Human Contracting Skeletal Muscle – Influence of Muscle Glycogen Content. FASEB, 15: 2748-2750.

17. Pilegaard H., C Keller, A Steensberg, J Wulff Helge, B Klarlund-Petersen, B Saltin, PD Neufer. 2002. Importance of glycogen content for the exercise-induced gene expression in human skeletal muscle. J. Physiol. 541: 261-271.

18. Wojtaszewski, J, M Mourtazkis, B Saltin, H Pilegaard. 2002. Progressive phosphorylation and activation of 5’ AMP activated protein kinase during prolonged low intensity exercise in humans. BBRC 298:309-316.

19. Nielsen JN, KJW Mustard, DA Graham, H Pilegaard, H Yu, LJ Goodyear, DG Hardie, EA Richter, and JFP Wojtaszewski. 2003. 5’AMP-activated protein kinase activity and subunit expression in exercise-trained human skeletal muscle. J. Appl Physiol 94(2):631-41.

20. Pilegaard, H, B Saltin, and PD Neufer. 2003. Short-term fasting and re-feeding on transcriptional regulation of metabolic genes in human skeletal muscle. Diabetes52: 657-662.

21. Pilegaard, H, B Saltin, and PD Neufer. 2003. Exercise induces transient transcriptional activation of the PGC-1 gene in human skeletal muscle. J Physiol. 546: 851-858.

22. Psilander N, R Damsgaard, H Pilegaard.2003. Resistance training alters the mRNA content of MRF’s and IGF-1 in human skeletal muscle. J Appl Physiol 95: 1038-44.

23. Nordsborg N, C Klarskov, P Krustrup, D Hudson, J Bangsbo, and H Pilegaard. 2003. Exercise-induced changes in gene expression related to metabolism and ion homeostasis before and after a high-intensity exercise training period. J Appl Physiol 95(3):1201-1206.

24. Hiscock N, C Fisher, H Pilegaard, B Klarlund Petersen. 2003. Vasuclar endothelial growth factor mRNA expression and a-v balance in response to prolonged, submaximal exercise in humans. Am J Physiol 285: H1759-1763.

25. Lundby C, H Pilegaard, G van Hall, M Sander, JAL Calbet, S Loft, and P Møller. 2003. Oxidative DNA damage and repair in skeletal muscle of humans exposed to hypoxia. Toxicology 5;192: 229-236.

26. Hildebrandt AL, H Pilegaard, PD Neufer. 2003. Differential Transcriptional Activation of Select Metabolic Genes in Response to Variations in Exercise Intensity and Duration in Red and White Skeletal Muscle. Am J Physiol 285: E1021-E1027.

27. Jensen L, H Pilegaard, Y Hellsten. 2004. Effect of acute exercise and exercise training on VEGF splice variants in human skeletal muscle. Am J Physiol 287:R397-R340.

28. Fischer CP, P Plomgaard, AK Hansen, H Pilegaard, B Saltin, BK Pedersen. 2004. Endurance training reduces the contraction-induced interleukin-6 mRNA expression in human skeletal muscle. Am J Physiol 287(6):E1189-1194.

29. Lundby C, Pilegaard H, can Hall G, Sander M, Calbet JA. 2004. Acclimatization to 4100 m does not change capillary density or mRNA expression of potential angiogenesis regulatory factors in human skeletal muscle. J Exp Biol 207(Pt 22): 3865-71.

30. Lindegaard B, AB Hansen, H Pilegaard, P Keller, J Gerstorf, BK Pedersen. 2004. Adipose tissue expression of IL-18 and HIV-associated lipodystrophy. AIDS 24:18(14):1956-1958.

31. Jørgensen SB, JFP Wojtaszewski, B Viollet, F Andreelli, Y Hellsten, P Scherling, S Vaulont, PD Neufer, EA Richter, H Pilegaard. 2005. Effect of α-AMPK knockout on exercise-induced gene activation in mouse skeletal muscle. FASEB J 19(9): 1146-8.

32. Pilegaard H, T Osada, Andersen L, JW Helge, B Saltin, PD Neufer. 2005. Influence of substrate availability on transcriptional regulation of metabolic genes in human skeletal muscle. Metabolism, 54(8):1048-55.

33. Wojtaszewski JFP, JB Birk, C Frøsig, M Holten, H Pilegaard and F Dela. 2005. 5'AMP activated protein kinase expression in human skeletal muscle: effects of strength training and type 2 diabetes. J Physiol 564: 563-73.

34. Nordsborg N, M Thomassen, C Lundby, H Pilegaard, J Bangsbo. 2005. Contraction-induced increases in Na+-K+-ATPase mRNA levels in human skeletal muscle are not amplified by activation of additional muscle mass. Am J Physiol 289: R84-91.

35. Lundby C, N Nordsborg, K Kusuhara, K Møller Kristensen, PD Neufer, H Pilegaard. 2005. Gene expression in human skeletal muscle: alternative normalization method and effect of repeated biopsies. Eur J Applied Physiol 95(4):351-60.

36. Lundby C, Gassmann M, Pilegaard H. 2006. Regular endurance training reduces the exercise induced HIF-1alpha and HIF-2alpha mRNA expression in human skeletal muscle in normoxic conditions. Eur J Appl Physiol 12: 1-7.

37. Mourtzakis M, Saltin B, Graham T, Pilegaard H. 2006. Carbohydrate Metabolism During Prolonged Exercise and Recovery: Interactions between Pyruvate Dehydrogenase, Fatty Acids and Amino Acids. J Appl Physiol 1822-30.

38. Carey A, Pedersen E, Southgate R, Bruce C, Pilegaard H, Hawley J, Pedersen BK, Febrario M. 2006. Discordant gene expression in skeletal muscle and adipose tissue of patients with Type 2 Diabetes: effect of IL-6 infusion. Diabetologica 49: 1000-1007.

39. Plomgaard P, Penkowa M, Pedersen BK, Saltin B, Pilegaard H. 2006. The mRNA expression profile of metabolic genes relative to MHC isoform pattern in human skeletal muscles. Am J Physiol 101: 817-825.

40. Roepstorff C, Thiele M, Hillig T, Pilegaard H, Hardie DG, Richter EA, Wojtaszewski JFP, Kiens B. 2006. Skeletal muscle α2AMPK activation during submaximal exercise is higher in men than in women. J Physiol 574: 125-138.

41. Silveira LR, Pilegaard H, Kusuhara K, Curi R, Hellsten Y. 2006. The effect of reactive oxygen species and antioxidants on basal and contraction-induced gene expression of PGC-1α, UCP3 and HKII in primary rat skeletal muscle cells. Biochim Biophys Acta 1763: 969-976.

42. Pilegaard H, Birk JB, Sacchetti M, Mourtzakis M, Hardie DG, Stewart G, Saltin B, Neufer PD, van Hall G, Wojtaszewski JFP. 2006. PDH-E1a dephosphorylation and activation in human skeletal muscle during exercise; effect of intralipid infusion Diabetes: 55: 3020-3027.

43. Leick L, B Lindegaard, D Berg, B Saltin, H Pilegaard. 2007. Adipose Tissue IL-18 mRNA and plasma IL-18 content – relation to obesity and HOMA index. Obesity Research 15: 356-363.

44. Kusuhara K, L Jensen, Y Hellsten, H Pilegaard. 2007. Ca2+ signaling in regulating PGC-1α, PDK4 and HKII mRNA expression studied in primary rat skeletal muscle cell culture. Biol Chem 388: 481-488.

45. Krabbe KS, AR Nielsen, R Krogh-Madsen, P Plomgaard, P Rasmussen, C Erikstrup, CP Fischer, B Lindegaard, AMW Petersen, S Taudorf, NH Secher, H Pilegaard, H Bruunsgaard and BK Pedersen. 2007. Brain-derived neurotrophic factor (BDNF) and type 2 diabetes. Reply to Lambert GW et al (letter). Diabetologia 50: 431–438.

46. Hellsten H, JJ Nielsen, J Lykkesfeldt, M Bruhn, L Silveira, H Pilegaard, J Bangsbo. 2007. Antioxidant supplementation enhances the exercise-induced increase in mitochondrial uncoupling protein 3 and endothelial nitric oxide synthase mRNA content in human skeletal muscle. Free Radic Biol Med 43: 353-361.

47. Robach P, G Cairo, C Gelfi, F Bernuzzi, H Pilegaard, A Viganò, P Santambrogio, P Cerretelli, JAL Calbet, S Moutereau, and C Lundby. 2007. Strong iron demand during hypoxia-induced erythropoiesis is associated with down-regulation of iron-related proteins and myoglobin in human skeletal muscle. Blood 109: 4724-4731.

48. Helge JW, NJ Rehrer, Pilegaard H, P Manning, SJ Lucas, DE Gerrard, JD Cotter. 2007. Increased fat

oxidation and regulation of metabolic genes with ultraendurance exercise. Acta Physiol 191: 77-86.

49. Steensberg A, C Keller, Hillig T, C Frøsig, JFP Wojtaszewski, BK Pedersen, H Pilegaard, M Sander. 2007. Nitric oxide production is a proximal signaling event controlling exercise-induced mRNA expression in human skeletal muscle. FASEB J 21: 2683-2694.

50. AR Nielsen, R Mounier, P Plomgaard, OH Mortensen, M Penkowa, T Speerschneider, R Damsgaard, H Pilegaard, BK Pedersen. 2007. Expression of IL-15 in human skeletal muscle. J Physiol 584: 305-312.

51. Krabbe KS, Mortensen EL, Avlund K, Pilegaard H, Christiansen L, Pedersen AN, Schroll M, Jørgensen T , Pedersen B K, Bruunsgaard H. 2007. Genetic priming of a proinflammatory profile predicts low IQ in octogenarians, Neurobiology of Aging 30: 769-81.

52. Mortensen OH, Plomgaard P, Fischer CP, Hansen AK, Pilegaard H, Pedersen BK. 2007. PGC-1{beta} is Downregulated by Training in Human Skeletal Muscle: No Effect of Training Twice Every Second Day versus Once Daily on Expression of The PGC-1 Family, J Appl Physiol 103: 1536-1542.

53. Klein DK, H Pilegaard, JT Treebak, TE Jensen, B Viollet, P Schjerling, EA Richter, JFP Wojtaszewski. 2007. Lack of a2 5´AMP Activated Protein Kinase enhances Pyruvate Dehydrogenase activity during in vivo exercise. Am J Physiol 293(5): E1242-1249.

*54. Leick L, Wojtaszewski JFP, Johansen ST, Kiilerich K, Comes G, Hellsten Y, Hidalgo J, Pilegaard H. 2008. PGC-1{alpha} is not mandatory for exercise- and training-induced adaptive gene responsens in mouse skeletal muscle. Am J Physiol Endocrinol Metab 294: E463-74.

55. Kiilerich K, Birk JB, Damsgaard R, Wojtaszewski JFP, Pilegaard H. 2008. Regulation of PDH in human arm and leg muscles at rest and during intense exercise. Am J Physiol Endocrinol Metab 294: E36-42.

56. Lundby C, Hellsten Y, Jensen MB, Munch AS, Pilegaard H. 2008. Erythropoietin receptor in human skeletal muscle and the effects of acute and long term injections with recombinant human erythropoietin on the skeletal muscle. J Appl Physiol 104: 1154-60.

57. Pilegaard H & Richter EA. 2008. PGC-1{alpha}: important for exercise performance? J Appl Physiol 104: 1264-1265.

58. Mortensen SP, González-Alons J Bune L, Saltin B, Pilegaard H, Hellsten Y. 2009. ATP induced vasodilatation and purinergic receptors in the human leg: roles of nitric oxide, prostaglandins and adenosine. Am J Physiol 296:R1140-8.

59. Coffey VG, Pilegaard H, Garnham AP, O’Brien BJ, Hawley1 JA. 2009. Consecutive bouts of diverse contractile activity alter acute responses in human skeletal muscle. J Appl Physiol 106: 1187-97.

60. Treebak JT, Frøsig C, Pehmøller C, Chen S, Maarbjerg SJ, Brandt N, MacKintosh C, Zierath JR, Hardie DG, Kiens B, Richter EA, Pilegaard H, Wojtaszewski JFP. 2009. Potential role of TBC1D4 in enhanced post-exercise insulin action in human skeletal muscle. Diabetologia 52: 891-900.

61. Leick L, Hellsten Y, Fentz J, Lyngby SS, Wojtaszewski JF, Hidalgo J, Pilegaard H. 2009. PGC-1{alpha} mediates exercise-induced skeletal muscle VEGF expression in mice. Am J Physiol 297: E92-103.

62. Adhihetty PJ, Uguccioni G, Leick L, Hidalgo J, Pilegaard H, Hood DA. 2009. Am J Physiol. The role of PGC-1{alpha} on mitochondrial function and apoptotic susceptibility in muscle. Am J Physiol 297: C217-C225.

63. Leick L, Plomgaard P, Wojtaszewski JFP, Pilegaard H. 2010. Exercise induces mRNA of oxidative proteins in human skeletal muscle late in recovery. Scan J Sports Medicine, 20: 593-599.

64. Rasmussen P, Brassard P, Adser H, Pedersen MV, Leick L, Hart, E, Secher NH, Pedersen BK, Pilegaard H. 2009. Evidence for a release of BDNF from the brain during exercise. Experimental Biology 94(10):1062-1069.

65. Hidalgo J, Florit S, Giralt M, Ferrer B, Keller C, Pilegaard H. 2009. Transgenic mice with astrocyte-targeted production of interleukin-6 are resistant to high-fat diet-induced increases in body weight and body fat. Brain Behav Immun 24(1): 119-126.

66. Thaning P, Bune LT, Hellsten Y, Pilegaard H, Saltin B, Rosenmeier JB. 2009. Attenuated Purinergic Receptor Function in Patients with Type 2 Diabetes. Diabetes 59(1): 182-189.

*67. Kiilerich K, Gudmundsson M, Birk JB, Lundby C, Taudorf S, Plomgaard P, Saltin B, Pedersen PA, Wojtaszewski JFP, Pilegaard H. 2010. Low Muscle Glycogen and Elevated Plasma FFA Modify but does not Prevent Exercise-induced PDH Activation in Human Skeletal Muscle. Diabetes 59(1): 26-32.

68. Nordsborg N, Kusuhara K, Hellsten Y, Lyndby S, Lundby C, Madsen K, Pilegaard H. 2010. Contraction induced changes in skeletal muscle Na(+), K(+) pump mRNA expression - importance of exercise intensity and Ca(2+) mediated signalling. Acta Physiol, 198(4): 487-498.

69. Leick L, Lyngby SS, Wojtaszewski JFP, Pilegaard H. 2010. PGC-1alpha is required for training-induced prevention of age-associated decline in mitochondrial enzymes in mouse skeletal muscle. Exp Gerontol 45: 336-342.

70. Seifert T, Brassard P, Wissenberg M, Rasmussen P, Nordby P, Stallknecht B, Adser H, Jakobsen AH, Pilegaard H, Nielsen HB, Secher NH. 2010. Endurance training enhances BDNF release from the human brain. Am J Physiol 298: R372-R377.

71. Nordsborg N, Lundby C, Leick L, Pilegaard H. 2010. Relative workload determines exercise-induced increases in PGC-1alpha mRNA. Med & Science in Sports & Exercise 42(8): 1477-1484.

72. Leick L, Fenz J, Biensø, RS, Knudsen JK, Jeppesen J, Kiens B, Wojtaszewski JFP, Pilegaard H. 2010. PGC-1{alpha} is required for AICAR induced expression of GLUT4 and mitochondrial proteins in mouse skeletal muscle. Am J Physiol 299: E456-465.

73. Kiilerich K, Adser H, Jakobsen AH, Pedersen PA, Hardie DG, Wojtaszewski JFP, Pilegaard H. 2010. PGC-1a increases PDH content but does not change the acute regulation of PDH in mouse skeletal muscle. Am J Physiol, 299(5):R1350-1359.

74. Olesen J, K Kiilerich, H Pilegaard. 2010. PGC-1alpha-mediated adaptations in skeletal muscle. Pflügers Arch 460: 153-162.

75. Høeg LD, Sjøberg KA, Jeppesen J, Jensen TE, Frøsig C, Birk JB, Bisiani B, Hiscock N, Pilegaard H, Wojtaszewski JF, Richter EA, Kiens B. 2011. Lipid-induced insulin resistance affects women less than men and is not accompanied by inflammation or impaired proximal insulin signaling. Diabetes 60(1): 64-73.

76. Rasmussen P, Vedel JC, Olesen J, Adser H, Pedersen MV, Hart E, Secher NH, Pilegaard H. 2011. In humans IL-6 is released from the brain during and after exercise and paralleled by enhanced IL-6 mRNA expression in the hippocampus of mice. Acta Physiol 201(4):475-82.

77. Pedersen L, Pilegaard H, Hansen J, Brandt C, Adser H, Hidalgo J, Olesen J, Pedersen BK, Hojman P. 2011. Exercise-induced liver CXCL-1 expression is linked to muscle derived IL-6 expression. J Physiol 589: 1409-20.

78. Adser H, Wojtaszewski JF, Jakobsen AH, Kiilerich K, Hidalgo J, Pilegaard H. 2011. Interleukin-6 modifies mRNA expression in mouse skeletal muscle. Acta Physiol 202: 165-173.

79. Iversen N, Krustrup P, Rasmussen HN, Rasmussen UF, Saltin B, Pilegaard H. 2011. Mitochondrial biogenesis and angiogenesis in skeletal muscle of the elderly. Exp Gerontol 46(8):670-678.

80. Broholm C, Laye MJ, Brandt C, Vadalasetty R, Pilegaard H, Pedersen BK, Scheele C. 2011. LIF is a contraction-induced myokine stimulating human myocyte proliferation. J Appl Physiol 111(1): 251-259.

81. Kiilerich K, Ringholm S, Biensø RS, Fisher JP, Iversen N, Van Hall G, Wojtaszewski JF, Saltin B, Lundby C, Calbet JA, Pilegaard H. 2011. Exercise-induced Pyruvate Dehydrogenase Activation Is Not Affected by Seven Days of Bed Rest. J Appl Physiol 111(3):751-757.

82. Ingwersen MS, Kristensen M, Pilegaard H, Wojtaszewski JF, Richter EA, Juel C. 2011. Na,K ATPase Activity in Mouse Muscle is Regulated by AMPK and PGC-1α. J Membr Biol 242(1): 1-10.

83. Ma T, Liaset B, Hao Q, Petersen RK, Fjære E, Ngo HT, Lillefosse HH, Ringholm S, Sonne SB, Treebak JT, Pilegaard H, Frøyland L, Kristiansen K, Madsen L. 2011. Sucrose counteracts the anti-inflammatory effect of fish oil in adipose tissue and increases obesity development in mice. PLoS One. 6(6):e21647.

84. Ringholm S, Biensø RS, Kiilerich K, Guadalupe-Grau A, Aachmann-Andersen NJ, Saltin B, Plomgaard P, Lundby C, Wojtaszewski JF, Calbet JA, Pilegaard H. 2011. Bed rest reduces metabolic protein content and abolishes exercise-induced mRNA responses in human skeletal muscle. Am J Physiol 301(4):E649-E658.

85. Haase TN, Jørgensen SR, Leick L, Biensø RS, Kiilerich K, Johansen S, Nielsen MM, Wojtaszewski JF, Hidalgo J, Pedersen PA, Pilegaard H. 2011. Role of PGC-1{alpha} in exercise and fasting induced adaptations in mouse liver. Am J Physiol 301(5):R1501-9.

86. Brandt C, Jakobsen A, Adser H, Olesen J, Iversen N, Kristensen J, Hojman P, Wojtaszewski J, Hidalgo J, Pilegaard H. 2011. IL-6 regulates exercise and training-induced adaptations in subcutaneous adipose tissue in mice. Acta Physiol 205(2):224-235.

87. Yfanti C, Fischer CP, Nielsen S, Akerström T, Nielsen AR, Veskoukis AS, Kouretas D, Lykkesfeldt J, Pilegaard H, Pedersen BK. 2012. Role of Vitamin C and E Supplementation on IL-6 in response to Training. J Appl Physiol. 112(6):990-1000.

88. Christensen B, Lundby C, Jessen N, Nielsen TS, Vestergaard PF, Møller N, Pilegaard H, Pedersen SB, Kopchick JJ, Jørgensen JO. 2012. Evaluation of functional erythropoietin receptor status in skeletal muscle in vivo: acute and prolonged studies in healthy human subjects. PLoS One. 2012;7(2):e31857.

89. Olesen J, Larsson S, Iversen N, Yousafzai S, Hellsten Y, Pilegaard H. 2012. Skeletal Muscle PGC-1α Is Required for Maintaining an Acute LPS-Induced TNFα Response. PLoS One. 2012;7(2):e32222.

90. Biensø RS, Ringholm S, Kiilerich K, Aachmann-Andersen NJ, Krogh-Madsen R, Guerra B, Plomgaard P, van Hall G, Treebak JT, Saltin B, Lundby C, Calbet JA, Pilegaard H, Wojtaszewski JF. 2012. GLUT4 and Glycogen Synthase Are Key Players in Bed Rest-Induced Insulin Resistance. Diabetes 61(5):1090-1099.

91. Sjøgaard G, Zebis MK, Kiilerich K, Saltin B, Pilegaard H. 2013. Exercise Training and Work Task Induced Metabolic and Stress-Related mRNA and Protein Responses in Myalgic Muscles. Journal of Biomed Res Int, 984523.

92. Mänttäri S, Ørtenblad N, Madsen K, Pilegaard H. 2013. Both short intense and prolonged moderate in vitro stimulation reduce the mRNA expression of calcium-regulatory proteins in rat skeletal muscle. Mol Cell Biochem, 373(1-2):171-178.

93. Buhl M, Bosnjak E, Vendelbo MH, Gjedsted J, Nielsen RR, K-Hafstrøm T, Vestergaard ET, Jessen N, Tønnesen E, Møller AB, Pedersen SB, Pilegaard H, Biensø RS, Jørgensen JO, Møller N. 2013. Effects of

Locally Administered Lipopolysaccharide on Glucose, Lipid, and Protein Metabolism in the Placebo-Controlled, Bilaterally Infused Human Leg. J Clin Endocrinol Metab 98(5): 2090-2099.

94. Abildgaard J, Pedersen AT, Green CJ, Harder-Lauridsen NM, Solomon TP, Thomsen C, Juul A, Pedersen M, Pedersen JT, Mortensen OH, Pilegaard H, Pedersen BK, Lindegaard B. 2013. Menopause is associated with decreased whole body fat oxidation during exercise. Am J Physiol 304(11): E1227-E1236.

95. Mortensen B, Hingst JR, Frederiksen N, Hansen RWW, Christiansen CS, Iversen N, Friedrichsen M, Birk JB, Pilegaard H, Hellsten Y, Vaag A, Wojtaszewski JFP. 2013. Effect of birth weight and 12 weeks of exercise training on exercise-induced AMPK signaling in human skeletal muscle. Am J Physiology 304(12):E1379-1390.

96. Ringholm S, Knudsen JG, Leick L, Lundgaard A, Nielsen MM, Pilegaard H. 2013. PGC-1alpha is required for exercise- and exercise training-induced UCP1 up-regulation in mouse white adipose tissue. PLos One, 22;8(5):e64123.

97. Hoier B, Prats C, Qvortrup K, Pilegaard H, Bangsbo J, and Hellsten Y. 2013. ubcellular localization and secretion of vascular endothelial growth factor in human skeletal muscle at rest and after exercise. FASEB J 27(9): 3496-3504.

98. Lindegaard B, Matthews VB, Brandt C, Hojman P, Allen TL, Estevez E, Watt MJ, Bruce CR, Mortensen O, Syberg S, Rudnicka C, Abildgaard J, Pilegaard H, Hidalgo J, Ditlevsen S, Alsted TJ, Madsen AN, Pedersen BK, Febbraio MA. 2013. Interleukin-18 activates skeletal muscle AMPK and reduces weight gain and insulin resistance in mice. Diabetes 62(9): 3064-3074.

99. Bach E, Nielsen RR, Vendelbo MH, Møller AB, Jessen N, Buhl M, Hafstrøm TK, Holm L, Pedersen SB, Pilegaard H, Biensø RS, Jørgensen JO, Møller N. 2013. Direct effects of TNF-α on local fuel metabolism and cytokine levels in the placebo controlled bilaterally infused human leg; increased insulin sensitivity, increased net protein breakdown and increased IL-6 release. Diabetes.

100. Gliemann L, Schmidt JF, Olesen J, Biensø RS, Peronard SL, Grandjean SU, Mortensen SP, Nyberg M, Bangsbo J, Pilegaard H, Hellsten Y. 2013. Resveratrol Blunts the Positive Effects of Exercise Training on Cardiovascular Health in Aged Men. J Physiol 15 (591): 5047-5059.

101. Olesen J, Ringholm S, Nielsen MM, Brandt CT, Pedersen JT, Halling JF, Goodyear LJ, Pilegaard H. 2013. Role of PGC-1α in exercise training- and resveratrol-induced prevention of age-associated inflammation. Exp Gerontol 48(11):1274-84.

102. Brandauer J, Vienberg SG, Andersen MA, Ringholm S, Risis S, Larsen PS, Kristensen JM, Frøsig C, Leick L, Fentz J, Jørgensen SB, Kiens B, Wojtaszewski JF, Richter EA, Zierath JR, Goodyear LJ, Pilegaard H, Treebak JT. 2013. AMP-activated protein kinase regulates nicotinamide phosphoribosyl transferase expression in skeletal muscle. J Physiol 591: 5207-5220.

103. Ringholm S, Olesen J, Pedersen JT, Brandt CT, Halling JF, Hellsten Y, Prats C, Pilegaard H. 2013. Effect of lifelong resveratrol supplementation and exercise training on skeletal muscle oxidative capacity in aging mice; impact of PGC-1α. Exp Gerontol 48(11): 1311-1318. 104. Nellemann B, Vendelbo MH, Nielsen TS, Bak AM, Høgild M, Pedersen SB, Biensø RS, Pilegaard H, Møller N, Jessen N, Jørgensen JO. 2013. Growth hormone-induced insulin resistance in human subjects involves reduced pyruvate dehydrogenase activity. Acta Physiol Oct 21. doi: 10.1111/apha.12183.

105. Dyar KA, Ciciliot S, Wright LE, Biensø RS, Tagliazucchi GM, Patel VR, Forcato M, Paz MI, Gudiksen A, Solagna F, Albiero M, Moretti I, Eckel-Mahan KL, Baldi P, Sassone-Corsi P, Rizzuto R, Bicciato S, Pilegaard H, Blaauw B, Schiaffino S. 2013. Muscle insulin sensitivity and glucose metabolism are controlled by the intrinsic muscle clock. Mol Metab 23;3(1): 29-41.

106. Knudsen JG, Murholm M, Carey AL, Biensø RS, Basse AL, Allen TL, Hidalgo J, Kingwell BA, Febbraio MA, Hansen JB, Pilegaard H. 2014. Role of IL-6 in exercise training- and cold-induced UCP1 expression in subcutaneous white adipose tissue. PLoS One 8;9(1).

107. Guerra B, Ponce-González JG, Morales-Alamo D, Guadalupe-Grau A, Kiilerich K, Fuentes T, Ringholm S, Biensø RS, Santana A, Lundby C, Pilegaard H, Calbet JA. 2014. Leptin signaling in skeletal muscle after bed rest in healthy humans. Eur J Appl Physiol 114(2): 345-57.

108. Olesen J, Gliemann L, Biensø R, Schmidt J, Hellsten Y, Pilegaard H. 2014. Exercise training, but not resveratrol, improves metabolic and inflammatory status in skeletal muscle of aged men. J Physiol 15; 592: 1873-86.

109. Biensø RS, Knudsen JG, Brandt N, Pedersen PA, Pilegaard H. 2015. Effects of IL-6 on pyruvate dehydrogenase regulation in mouse skeletal muscle. Pflugers Arch. 466(8): 1647-57.

110. Jordy AB, Serup AK, Karstoft K, Pilegaard H, Kiens B, Jeppesen J. 2014. Insulin Sensitivity is Independent of Lipid Binding Protein Trafficking at the Plasma Membrane in Human Skeletal Muscle: Effect of a 3-day, High-fat Diet. Am J Physiol Regul Integr Comp Physiol. 1;307(9): R1136-45.

111. Olesen J, Biensø RS, Meinertz S, van Hauen L, Rasmussen SM, Gliemann L, Plomgaard P, Pilegaard H. 2014. Impact of Training Status on LPS-induced Acute Inflammation in Humans. J Appl Physiol. 1;118(7): 818-29.

112. Biensø RS, Olesen J, van Hauen L, Meinertz S, Halling JF, Gliemann L, Plomgaard P, Pilegaard H. 2015. Exercise-induced AMPK and pyruvate dehydrogenase regulation is maintained during short-term low-grade inflammation. Pflugers Arch 467 (2): 341-350.

113. Knudsen JG, Biensø RS, Hassing HA, Jakobsen AH, Pilegaard H. 2015. Exercise-induced regulation of key factors in substrate choice and gluconeogenesis in mouse liver. Mol Cell Biochem 403: 209-17.

114. Knudsen JG, Bertholdt L, Joensen E, Lassen SB, Hidalgo H, Pilegaard H. 2015. Skeletal muscle interleukin-6 regulates metabolic factors in iWAT during HFD and exercise training. Obesity. 23(8): 1616-24.

115. Biensø RS, Olesen J, Gliemann L, Schmidt JF, Matzen MS, Wojtaszewski JF, Hellsten Y, Pilegaard H. Effects of Exercise Training on Regulation of Skeletal Muscle Glucose Metabolism in Elderly Men. J Gerontol A Biol Sci Med Sci. 2015 70(7): 866-72, 2015.

116. Fritzen AM, Lundsgaard AM, Jordy AB, Kellebjerg Poulsen S, Stender S, Pilegaard H, Astrup A, Meinert Larsen T, Wojtaszewski JF, Richter EA, Kiens B. 2015. New Nordic Diet induced weight loss is accompanied by changes in metabolism and AMPK signalling in adipose tissue. J Clin Endocrinol Metab 100(9):3509-19.

117. Fritzen AM, Lundsgaard AM, Jeppesen J, Christiansen ML, Biensø R, Dyck JR, Pilegaard H, Kiens B. 2015. 5´AMP activated protein kinase α2 controls substrate metabolism during post-exercise recovery via regulation of pyruvate dehydrogenase kinase 4. J Physiol. 1;593(21):4765-80.

118. Fentz J, Kjøbsted R, Kristensen CM, Hingst JR, Birk JB, Gudiksen A, Foretz M, Schjerling P, Viollet B, Pilegaard H, Wojtaszewski JF. 2015. AMPKα is essential for acute exercise-induced gene responses but not for exercise training-induced adaptations in mouse skeletal muscle. Am J Physiol Endocrinol Metab. 1;309(11):E900-14.

119. Edge J, Mündel T, Pilegaard H, Hawke E, Leikis M, Lopez-Villalobos N, Oliveira RS, Bishop DJ. 2015. Ammonium Chloride Ingestion Attenuates Exercise-Induced mRNA Levels in Human Muscle. PLoS One 10;10(12):e0141317.

120. Halling JF, Ringholm S, Nielsen MM, Overby P, Pilegaard H. 2016. PGC-1α regulates exercise-induced autophagy in mouse skeletal muscle. Physiological Reports 4(3). pii:e12698. doi:10.14814/phy2.12698.

121. Knudsen JG, Joensen E, Bertholdt L, Jessen H, van Hauen L, Hidalgo J, Pilegaard H. 2016 Skeletal muscle IL-6 and regulation of liver metabolism during high fat diet and exercise training. Physiological Reports. Physiol Rep 4(9). pii: e12788. doi: 10.14814/phy2.12788.

122. Gudiksen A, Schwartz CL, Bertholdt L, Joensen E, Knudsen JG, Pilegaard H. 2016. Lack of Skeletal Muscle IL-6 Affects Pyruvate Dehydrogenase Activity at Rest and during Prolonged Exercise. PLoS One 21;11(6):e0156460. doi: 10.1371/journal.pone.0156460.

123. Brandt N, Gunnarsson TP, Hostrup M, Tybirk J, Nybo L, Pilegaard H, Bangsbo J. 2016. Impact of adrenaline and metabolic stress on exercise-induced intracellular signaling and PGC-1α mRNA response in human skeletal muscle. Physiological Reports 4(14). pii: e12844. doi: 10.14814/phy2.12844.

124. Skovgaard C, Brandt N, Pilegaard H, Bangsbo J. 2016. Combined speed endurance and endurance exercise amplify the exercise-induced PGC-1α and PDK4 mRNA response in trained human muscle. Physiological Reports 4(14). pii: e12864. doi: 10.14814/phy2.12864.

125. Gudiksen A, Bertholdt L, Vingborg MB, Hansen HW, Ringholm S, Pilegaard H. 2017. Muscle interleukin 6 and fasting-induced PDH regulation in mouse skeletal muscle. Am J Physiol Endocrinol Metab 1;312(3):E204-E214.

126. Bertholdt L, Gudiksen A, Schwartz CL, Knudsen JG, Pilegaard H. 2017. Lack of skeletal muscle IL-6 influences hepatic glucose metabolism in mice during prolonged exercise. Am J Physiol Regul Integr Comp Physiol 1; 312(4): R626-R636.

127. Gudiksen A & Pilegaard H. 2017. PGC-1α and fasting-induced PDH regulation in mouse skeletal muscle. Physiological Reports 5(7). pii: e13222. doi: 10.14814/phy2.13222.

128. Halling JF, Ringholm S, Olesen J, Prats C, Pilegaard H. 2017. Exercise training protects against aging-induced mitochondrial fragmentation in mouse skeletal muscle in a PGC-1α dependent manner. Experimental Gerontology 96:1-6.

129. Gudiksen A, Bertholdt L, Stankiewicz T, Tybirk J, Plomgaard P, Bangsbo J, Pilegaard H. 2017. Effects of training status on PDH regulation in human skeletal muscle during exercise. Pflugers Arch. 469(12):1615-1630.

130. Kristensen CM, Brandt CT, Ringholm S, Pilegaard H. 2017. PGC-1α in aging and lifelong exercise training-mediated regulation of UPR in mouse liver. Exp Gerontol 98:124-133.

131. Brandt N, Nielsen L, Buch B, Gudiksen A, Ringholm S, Hellsten Y, Bangsbo J, Pilegaard H. 2017. Impact of β-adrenergic signaling in PGC-1α-mediated adaptations in mouse skeletal muscle. Am J Physiol 1;314(1): E1-E20.

132. Lundsgaard AM, Sjøberg KA, Høeg LD, Jeppesen J, Jordy AB, Serup AK, Fritzen AM, Pilegaard H, Myrmel LS, Madsen L, Wojtaszewski JFP, Richter EA, Kiens B. 2017. Opposite Regulation of Insulin Sensitivity by Dietary Lipid Versus Carbohydrate Excess. Diabetes 66: 2583-2595.

133. Brandt N, Dethlefsen MM, Bangsbo J, Pilegaard H. 2017. PGC-1α and exercise intensity dependent adaptations in mouse skeletal muscle. PLOS ONE. 19;12(10):e0185993. doi: 10.1371/ journal.pone. 0185993.

134. Bertholdt L, Gudiksen A, Stankiewicz T, Villesen I, Tybirk J, van Hall G, Bangsbo J, Plomgaard P, Pilegaard H. 2018. Impact of training state on fasting-induced regulation of adipose tissue metabolism in humans. J Appl Physiol 1;124(3):729-740. doi: 10.1152/japplphysiol.00664.2017.

135. Knudsen JG, Gudiksen A, Bertholdt L, Overby P, Villesen I, Schwartz CL, Pilegaard H. 2017. Skeletal muscle IL-6 regulates muscle substrate utilization and adipose tissue metabolism during recovery from an acute bout of exercise. PLoS One. 2017 Dec 18;12(12):e0189301. doi: 10.1371/journal.pone. 0189301. eCollection.

136. Rasmussen MK, Bertholdt L, Gudiksen A, Pilegaard H, Knudsen JG. 2018. Impact of fasting followed by short-term exposure to interleukin-6 on cytochrome P450 mRNA in mice. Toxicol Lett 5;282:93-99. doi: 10.1016/j.toxlet.2017.10.011.

137. Møller HD, Mohiyuddin M, Prada-Luengo I, Sailani MR, Halling JF, Plomgaard P, Maretty L, Hansen AJ, Snyder MP, Pilegaard H, Lam HYK, Regenberg B. 2018. Circular DNA elements of chromosomal origin are common in healthy human somatic tissue. Nat Commun 14;9(1):1069. doi: 10.1038/s41467-018-03369-8.

138. Brandt N, Gunnarsson TP, Bangsbo J, Pilegaard H. 2018. Exercise and exercise training-induced increase in autophagy markers in human skeletal muscle. Physiol Rep 6(7):e13651. doi: 10.14814/phy2.13651.

139. Fiorenza M, Gunnarsson TP, Hostrup M, Iaia FM, Schena F, Pilegaard H, Bangsbo J. 2018. Metabolic stress‒dependent regulation of the mitochondrial biogenic molecular response to high-intensity exercise in human skeletal muscle. J Physiol 96: 2823-2840 doi: 10.1113/JP275972.

140. Dethlefsen MM, Kristensen CM, Tøndering AS, Lassen SB, Ringholm S, Pilegaard H. 2018. Impact of liver PGC-1α on exercise and exercise training-induced regulation of hepatic autophagy and mitophagy in mice on HFF. Physiological Reports 6(13):e13731. doi: 10.14814/phy2.13731.

141. Kristensen CM, Olsen MA, Jessen H, Brandt N, Meldgaard JN, Pilegaard H. 2018. PGC-1α in exercise and fasting-induced regulation of hepatic UPR in mice. Plugers Arch 470(10):1431-1447.

142. Kristensen CM, Jessen H, Ringholm S, Pilegaard H. 2018. Muscle PGC-1α in exercise and fasting-induced regulation of hepatic UPR in mice. Acta Physiol :e13158. doi: 10.1111/apha.13158.

143. Hostrup M, Reitelseder S, Jessen S, Kalsen A, Nyberg M, Egelund J, Kreiberg M, Kristensen CM, Thomassen M, Pilegaard H, Backer V, Jacobson GA, Holm L, Bangsbo J. 2018. Beta2 -adrenoceptor

agonist salbutamol increases protein turnover rates and alters signalling in skeletal muscle after resistance exercise in young men. J Physiol 596(17):4121-4139.

144. Gudiksen A, Bertholdt L, Stankiewicz T, Villesen I, Bangsbo J, Plomgaard P, Pilegaard H. 2018. Training state and fasting-induced PDH regulation in human skeletal muscle. Pflugers Arch doi: 10.1007/s00424-018-2164-6.

145. Bertholdt L, Gudiksen A, Jessen H, Pilegaard H. 2018. Impact of skeletal muscle IL-6 on regulation of liver and adipose tissue metabolism during fasting. Pflugers Arch. 2018 Aug 1. doi: 10.1007/s00424-018-2185.

146. Kristensen CM, Dethlefsen MM, Tøndering AS, Lassen SB, Meldgaard JN, Ringholm S, Pilegaard H. 2018. PGC-1α in hepatic UPR during high-fat high-fructose diet and exercise training in mice. Physiol Rep Aug;6(15):e13819. doi: 10.14814/phy2.13819.

147. Dethlefsen MM, Halling JF, Møller HD, Plomgaard P, Regenberg B, Ringholm S, Pilegaard H. 2018. Regulation of apoptosis and autophagy in mouse and human skeletal muscle with aging and lifelong exercise training. Exp Gerontol 111:141-153. doi: 10.1016/j.exger.2018.07.011.

148. Dethlefsen MM, Bertholdt L, Gudiksen A Stankiewicz T, Bangsbo J, Plomgaard P, Pilegaard H. 2018. Training state and skeletal muscle autophagy in response to 36h of fasting. J Appl Physiol, 125(5):1609-1619. doi: 10.1152/japplphysiol.01146.2017

149. Brandt N, Kotowska D, Kristensen CM, Olesen J, Lützhøft DO, Halling JF, Hansen M, Abu Al-Soud W, Hansen L, Kiilerich P, Pilegaard H. 2018. The impact of exercise training and resveratrol supplementation on gut microbiota composition in high-fat diet fed mice. Physiol Rep 6(20):e13881. doi: 10.14814/phy2.13881.

150. Gunnarsson TP, Brandt N, Fiorenza M, Hostrup M, Pilegaard H, Bangsbo J. 2019. Inclusion of sprints in moderate intensity continuous training leads to muscle oxidative adaptations in trained individuals. Physiol Rep. 7(4):e13976. doi: 10.14814/phy2.13976.

151. Sailani MR, Halling JF, Møller HD, Lee H, Plomgaard P, Pilegaard H, Snyder MP, Regenberg B. 2019. Lifelong physical activity is associated with promoter hypomethylation of genes involved in metabolism, myogenesis, contractile properties and oxidative stress resistance in aged human skeletal muscle. Scientific Reports 1;9(1):3272. doi: 10.1038/s41598-018-37895-8.

152. Halling JF, Jessen H, Nøhr-Meldgaard J, Buch BT, Christensen NM, Gudiksen A, Ringholm S, Neufer PD, Prats C, Pilegaard H. PGC-1α regulates mitochondrial properties beyond biogenesis with aging and exercise training. Am J Physiol Endocrinol Metab. 2019 Sep 1;317(3):E513-E525. doi: 10.1152/ajpendo.00059.2019

153. Høgild ML, Gudiksen A, Pilegaard H, Stødkilde-Jørgensen H, Pedersen SB, Møller N, Jørgensen JOL, Jessen N. 2019. Redundancy in regulation of lipid accumulation in skeletal muscle during prolonged fasting in obese men. Physiol Rep. Nov;7(21):e14285. doi: 10.14814/phy2.14285.

154. Thøgersen R, Kristensen CM, Olsen MA, Bertram HC, Pilegaard H, Rasmussen MK. 2020. Hepatic PGC-1α is not essential for fasting-induced cytochrome p450 regulation in mouse liver. Biochem Pharmacol. 29;172:113736. doi: 10.1016/j.bcp.2019.113736.

155. Bertholdt L, Gudiksen A, Ringholm S, Pilegaard H. 2020. Impact of skeletal muscle IL-6 on subcutaneous and visceral adipose tissue metabolism immediately after high- and moderate-intensity exercises. Pflugers Arch. 472(2):217-233; doi: 10.1007/s00424-019-02332-w.

156. Buch BT, Halling JF, Ringholm S, Gudiksen A, Kjøbsted R, Olsen MA, Wojtaszewski JFP, Pilegaard H. 2020. Colchicine Treatment Impairs Skeletal Muscle Mitochondrial Function and Insulin Sensitivity in an Age-Specific Manner. FASEB J. 34(6):8653-8670. doi: 10.1096/fj.201903113.

157. Hingst JR, Kjøbsted R, Birk JB, Jørgensen NO, Larsen MR, Kido K, Larsen JK, Kjeldsen SAS, Fentz J, Frøsig C, Holm S, Fritzen AM, Dohlmann TL, Larsen S, Foretz M, Viollet B, Schjerling P, Overby P, Halling JF, Pilegaard H, Helsten Y, Wojtaszewski JFP. 2020. Inducible Deletion of Skeletal Muscle AMPKα Reveals That AMPK Is Required for Nucleotide Balance but Dispensable for Muscle Glucose Uptake and Fat Oxidation During Exercise. Mol Metab 3:101028. doi: 10.1016/j.molmet.2020.101028.

158. Terzic D, Zois NE, Hunter I, Christoffersen C, Plomgaard P, Olsen LH, Ringholm S, Pilegaard H, Goetze JP. (2020). Effect of insulin on natriuretic peptide gene expression in porcine heart. Peptides. 131:170370. doi: 10.1016/j.peptides.2020.170370. Epub 2020 Jul 11.PMID: 32663503.

159. Hjelholt AJ, Charidemou E, Griffin JL, Pedersen SB, Gudiksen A, Pilegaard H, Jessen N, Møller N, Jørgensen JOL. (2020). Insulin resistance induced by growth hormone is linked to lipolysis and associated with suppressed pyruvate dehydrogenase activity in skeletal muscle: a 2 × 2 factorial, randomised, crossover study in human individuals. Diabetologia. 2020 Sep 18. doi: 10.1007/s00125-020-05262-w. Online ahead of print.PMID: 32945898.

Proceedings, reviews and book chapters

1. Pilegaard H and PD Neufer. 2004. Transcriptional regulation of PDK4 in skeletal muscle during and after exercise. Biochemistry of Exercise. Proc Nutr Soc 63:221-226.

2. Juel, C and H Pilegaard. 1998. Lactate exchange and pH regulation in skeletal muscle. In: Biochemistry of Exercise. Chapter 15: 185-200.

3. Joseph AM, H Pilegaard, A Litvintsev, L Leick and DA Hood. 2006. Control of Gene Expression and Mitochondrial Biogenesis in the Muscular Adaptation to Endurance Exercise. Essays in Biochemistry.

4. Wojtaszewski JFP, Pilegaard H, Dela F. 2007. Resistance exercise training in management of Type 2 diabetes (chapter 12). In: Physical activity and Type 2 diabetes. Therapeutic effects and mediators og action. Human Kinetics. Editors: Hawley J & Zierath J.

5. Halling, JF and Pilegaard H. 2017. Autophagy-dependent beneficial effects of exercise. Biology of Exercise. Editors J Zierath, J Hawley and M Joyner. Cold Spring Harb Perspect Med. Mar 7. pii: a029777. doi: 10.1101/cshperspect.a029777.

6. Halling JF, Pilegaard H. PGC-1α-mediated Regulation of Mitochondrial Function and Physiological Implications. Appl Physiol Nutr Metab. 2020 Jun 9. doi: 10.1139/apnm-2020-0005.

Menneskets Fysiologi (Human Physiology)

Cellulær Integrativ Fysiologi (Cellular Integrative Physiology)

Zoofysiologi (Zoophysiology)

Genteknologi (Gene technology)

Komparativ Anatomi (Comparative Anatomy)